Desmond Miles was a member of the Assassins and a descendant of numerous familial lines that had sworn an allegiance to the Assassins, including individuals such as Aquilus, Altaïr Ibn-La'Ahad, Ezio Auditore, Edward Kenway, and Ratonhnhaké:ton. He was also the son of the Mentor William Miles and unknowingly the father of Elijah, a young Sage.