Mira Nova is the Princess of the planet Tangea and also a Space Ranger as Buzz's co-pilot and second-in-command.

Being a princess, Mira is heir to Throne of Tangea. She became a space ranger against her father's wishes because she was inspired by how Star Command saved her planet. Her race is very aristocratic, something she dislikes. Tangea is home to two classes of people, Grounders, who live on the surface, and Mira's race, who lives in the sky and looks down upon the Grounders, both literally and figuratively. Mira Nova went against the royal family by dating a Grounder one year before she enlisted in Star Command, and also makes it clear that as a future leader of Tangea she wishes to stop the treatment of Grounders as outsiders. Unlike many females in the series, Mira Nova is one of the few women not dazzled by the bravado of Buzz Lightyear and can work with him as an equal partner.

In Buzz Lightyear of Star Command: The Adventure Begins, a direct-to-video film that serves as a pilot to the series, Mira is introduced as a rookie space ranger. Buzz is called by Commander Nebula to observe her doing very well in a training session at Star Command, showing herself to be highly skilled, and so she is assigned to Buzz Lightyear as his new partner. However, she is rebuffed by Buzz, who lost his previous partner, Warp Darkmatter during an investigation into 3 missing LGMs.

When Nebula is planning a full-scale assault on Planet Z, home to Zurg, Mira proposes an alternate plan. The plan called for a single ranger to slip past Zurg's defenses in the Alpha One, a small prototypical spacecraft. Though Buzz agrees, the plan is laughed off by Nebula, who accuses the two of being show-offs. Later on, Buzz and Mira both attempt to steal the ship, but Mira is successful, due to the usage of her mental powers. She flies off, pursued by Buzz in his spacecraft.

Because they leave, they avoid being victims of Zurg's plan: to use the Uni-Mind, a sphere that controls the LGMs telepathic bond, to control everyone in Star Command. Mira and Buzz, along with XR and Booster, who had stowed away in Buzz's ship, discover the plan and narrowly avoid being destroyed. Buzz insists on facing Zurg alone, but Mira and the others turned back after hearing Buzz admit in his "Misson Log" that he was wrong to try to work alone and are able to rescue Buzz from becoming Zurg's accomplice. Though Zurg escapes and Buzz's ex-partner is arrested, Mira is able to repair the uni-mind so that the mind-controlled rangers are freed. Afterward, she joins Buzz, XR, and Booster informing "Team Lightyear".

The TV series follows the adventures of Team Lightyear, which includes Mira, in their battle against Emperor Zurg. Mira is shown to act as a co-pilot and second-in-command to Buzz. She is also shown to possess the most common sense of anyone on the team. Mira is also shown to have issues with her father, who wants her to quit being a space ranger. He often insults Mira's teammates and comments on Mira's lack of practice with her mental powers.