One of the weapons of the Mysterian invaders, Moguera was unleashed in Japan, where it destroyed a village by triggering severe earthquakes while underground. Moguera soon surfaced and began rampaging through the countryside before the JSDF blew up a bridge as the machine was crossing it, destroying Moguera. The Mysterians would deploy a second Moguera to defend their dome when the Earth Defense Force began attacking it with Markalite Cannons. The second Moguera accidentally burrowed too close to a Markalite Cannon, which toppled onto it and destroyed it.

(Can't believe I forgot Moguera came first before Varan, what am I a fake fan!? SHIT!)